What does this Barbary Macaque facial expression mean?

What does this Barbary Macaque facial expression mean?

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A girl pretended having a piece of food in her hand and pointed it to a Barbary Macaque in Gibraltar. As a response he made this facial expression to her. What does this mean?

It looks like it is trying too threat.


Individual differences in scanpaths correspond with serotonin transporter genotype and behavioral phenotype in rhesus monkeys (Macaca mulatta)

That is a threat face. Barbary macaque threat faces often appear with a brow raise, lowered head, and an o-shaped mouth, sometimes with and sometimes without a vocalization. Given the context you described it is not surprising the girl received a threat. *Based on personal research experience

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