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What does this Barbary Macaque facial expression mean?

What does this Barbary Macaque facial expression mean?


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A girl pretended having a piece of food in her hand and pointed it to a Barbary Macaque in Gibraltar. As a response he made this facial expression to her. What does this mean?


It looks like it is trying too threat.

Source:

Individual differences in scanpaths correspond with serotonin transporter genotype and behavioral phenotype in rhesus monkeys (Macaca mulatta)


That is a threat face. Barbary macaque threat faces often appear with a brow raise, lowered head, and an o-shaped mouth, sometimes with and sometimes without a vocalization. Given the context you described it is not surprising the girl received a threat. *Based on personal research experience


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